Barry Conn (NSW) & Kipiro Damas (LAE).
Copyright held by the authors, Royal Botanic Gardens and Domain Trust, and Papua New Guinea National Herbarium

Levieria squarrosa Perkins

Botanische Jahrbucher für Systematik, Pflanzengeschichte und Pflanzengeographie Vol. 52: 196 (1915)

Other Literature: W .Philipson, Flora Malesiana Series 1 Vol. 10:

Field Characters: Small trees 10-20 m high. Bole cylindrical 25 cm diam.; almost straight bole 15 m long; buttresses absent; spines absent; aerial roots absent; stilt roots absent. Bark grey or brown, almost smooth, slightly pustular; lenticels irregular; subrhytidome yellow or green; bark thickness <25 mm thick, 5.0 mm thick; blaze consisting of 2 layers; strongly aromatic; pleasant; outer blaze brown or yellow, with stripes, granular without splinters; inner blaze red or brown, with stripes, fibrous; exudate present, colourless, spotty, changing colour on exposure to air, to brown or orange, not sticky. Terminal buds not enclosed by leaves; complex hairs absent; stinging hairs absent; mature twig without hairs.

Leaves: Spaced along branches <internodes readily visible>. Leaves spiral, simple; petiole present 5-10 mm long, not winged, attached to base of leaf blade <not peltate>, not swollen; lamina broadest at or near middle, 4.0-9.0 (-11.5) cm long, 1.2-3.3 (-4.8) cm wide; lamina symmetric, margin shortly crenate usually indistinct, with 1 or 2, less often up to 10 small blunt teeth present or entire, not dissected or lobed, apex acuminate or obtuse, venation pinnate <secondary veins arising from the midrib along its length>, secondary veins open <spaced far apart to easily see tertiary veins>, not prominent, but visible, intramarginal veins absent; lower surface pale green or green, upper surface dark green subglossy, hairs absent; oil dots absent; domatia absent; stipules absent.

Flowers: Inflorescence axillary <from between a leaf and branch> and terminal <at branch ends>. Flowers arising from a single point and flowers on a branched axis flower clusters on long stalks (peduncles); flowers bisexual, not stalked, with many planes of symmetry <actinomorphic> perianth present, pale yellow; ovary superior <seated above petals and sepals; hypogenous>, carpels separate (when more than one) <apocarpous>, locules 16-17.

Fruits: Fruits arising from single point surrounded by a yellow to yellow-brown receptacle or fruits arranged on branched axis fruiting clusters on long stalks. Fruit 8-10 mm long, 5.0 mm diam., fruit green or orange, not spiny, fleshy, simple of separate carpels of the one flower, indehiscent, drupe.

Distribution: Eastern Highlands, East Sepik, Morobe, Western Highlands, Southern Highlands, Central & West Sepik.

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